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Eogastropoda |
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Eogastropoda |
| GASTROPODA | Euomphalida Cladogram |
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Gastropoda (crown group) |--EOGASTROPODA | |-?OPHILETOIDEA | `--+--MACLURITOIDEA | `--+--EUOMPHALIDA | |--PLATYCERATOIDEA | |--NEOMPHALIDA | `--Patellogastropoda `--Orthogastropoda |
|
<== o EUOMPHALINA de Koninck, 1881
|-- Prohelicotoma
|-- Ophileta subraplana
`--+-- Ophileta complanata
`--+-- Lecanospira compacta
|-- Lecanospira nereine
`--+-- Barnesella ?lecanospiroides
|--o Ecculiomphalus Portlock, 1843
| |-- Malayaspira rugosa
| `--+-- MACLURITOIDEA Fischer, 1885
| `--+-- Malayaspira hintzei
| |--+--+-- Russospira harrisae
| | | `-- Ecculiomphalus bucklandi
| | `--o Helicotomidae Wenz, 1938
| | |?- Linsleyella Rohr, 1980
| | |-- Lophonema peccatonica Ulrich, in Purdue & Miser, 1916
| | |--+-- Polehemia taneyensis
| | | `-- Walcottoma frydai
| | `--+--+--o Ophiletina Ulrich & Scofield, 1897
| | | | |-- Oriostoma bromidensis
| | | | `--+-- O. sublaxa
| | | | `-- O. angularis
| | | `--o Helicotoma Salter, 1859
| | | |-- H. tennesseensis
| | | |--+-- H. planulata
| | | | `--+-- H. robinsoni
| | | | `-- H. blodgetti
| | | `--o Palaeomphalus Koken, 1925
| | | |-- P. ?gradatus
| | | `-- Helicotoma? sp. [Girvan sp.]
| | `--+--o Anomphalidae Wenz, 1938
| | | `--o Trochomphalus Koken, 1925
| | | |-- Trochomphalus ?dimidiatus
| | | `--+-- Straporillina sp. cf. S. circe (sensu Rohr, 1988)
| | | `--+-- Grantlandispira christei
| | | `--o Pycnomphalus Lindström, 1884
| | | |-- P. acutus
| | | `--+-- P. obesus
| | | `-- Turbocheilus immaturum
| | `--o Euomphalidae de Koninck, 1881
| | |?- Micromphalus Knight, 1945
| | |--o Boucotspira Rohr, 1980
| | | |-- B. cariniferus [Euomphalopterus cariniferus]
| | | `-- B. ?ordovicicus [Euomphalopterus ?ordovicicus]
| | |--o Euomphalopterus Roemer, 1876
| | | |-- E. subcarinatus
| | | |-- E. alatus
| | | `--+-- E. togatus
| | | `--+--+-- E. praetextus
| | | | `-- E. undulans
| | | `--o Spinicharybdis Rohr & Packard, 1982
| | | |-- S. frenatus [Euomphalopterus frenatus]
| | | `-- S. wilsoni
| | |--o Poleumita Clarke & Ruedemann, 1903
| | | |-- P. alata
| | | `--+-- P. discors
| | | |-- P. granulosa
| | | |-- Centrifugus planorbis Bronn, 1834
| | | |--o Nodonema Linsley, 1968
| | | | |-- N. rugosa [Poleumita rugosa]
| | | | `-- N. octavia [Poleumita octavia]
| | | `--+-- Euomphalus Sowerby, 1814
| | | `-- Straparollus de Montfort, 1810
| | `--o Pseudophoridae Miller, 1889
| | |?- Elasmonema Fischer, 1885
| | |-- Streptotrochus lamellosus [Boucotspira [in partim]]
| | `--+-- Streptotrochus? visbeyensis [Boucotspira [in partim]]
| | `--+--o Streptotrochus Perner, 1903
| | | |-- S. incisus
| | | `--+-- S. lundgreni
| | | `-- Hystricoceras astraciformis Jahn, 1894
| | `--o Discordichilus Cossmann, 1918
| | |-- Streptotrochus sp. aff. S. incisus
| | `--+-- D. gotlandicus [Siluriphorus gotlandicus]
| | |-- D. undulans [Siluriphorus undulans]
| | |--+--+-- D. dalli
| | | | `-- D. mollis
| | | `--o Pseudotectus Perner, 1903
| | | |-- P. kolmodoni [Discordichilus kolmodoni]
| | | `-- P. comes
| | `--o Pseudophorus Meek, 1873
| | |-- P. stuxbergi
| | `-- P. profundus
| `--o Lesueurillidae Wagner, 2002
| |-- "Lesueurilla" declivis
| |--o Eccyliopterus Remelé, 1888
| | |-- E. regularis
| | `--+-- E. ?princeps
| | `--+-- E. alatus
| | `-- E. owenanus
| `--+-- "Lesueurilla" prima
| |--o Lesueurilla Koken, 1898
| | |-- "Eccyliopterus" louderbacki
| | `--+-- Lesueurilla infundibula
| | `-- "Eccyliopterus" beloitensis
| |--o Mestoronema Wagner, 2002
| | |-- "Lesueurilla" marginalis
| | `--+-- "Lesueurilla" scotica
| | `-- "Lesueurilla" bipatellare
| `--o Pararaphistoma Vostokova, 1955
| |-- Climacoraphistoma vaginati
| |-- Climacoraphistoma damesi
| `--+-- Pararaphistoma qualteriana
| `-- Pararaphistoma schmidti
`--o Asgardaspira Wagner, 2002
|-- Barnesella measuresae
`--+-- Lytospira yochelsoni
`--+--o Lytospira Koken, 1896
| |-- L. angelini
| `-- L. subrotunda
`--+--+-- Lytospira gerrula
| `-- Lytospira ?norvegica
`--o EUOMPHALOIDEA de Koninck, 1881
`--o Raphistomatidae Koken, 1896
|-- Ceratopea canadensis
|-- Orospira bigranosa Butts, 1926
|--o Bridgeites Flower, 1968a
| |-- B. planodorsalis
| `--+-- B. supraconvexa
| `-- B. ?disjuncta
`--+-- Ceratopea ?laurentia
|--+-- Ceratopea pygmaea
| `-- Ceratopea unguis
`--+-- Pararaphisoma lemoni
`--+-- Helicotoma medffraensis Salter, 1859
`--+--o Scalites Emmons, 1842
| |-- "Palaeomphalus" giganetus Koken, 1925
| `--+-- Scalites katoi
| `--+--+-- Raphistoma tellerensis
| | `-- Raphistoma peracuta
| `--+-- Scalites angulatus
| `--o Raphistoma Hall, 1847
| |-- Raphistoma striata
| `-- Helicotoma gubanovi
`--o Holopeidae Wenz, 1938
|?- Umbospira Perner, 1903
|--o Raphistomina Ulrich & Scofield, 1897
| |-- R. lapicida
| |-- R. aperta
| |-- R. fissurata
| `-- R. rugata
`--+-- Pachystrophia devexa
|--+-- Pachystrophia contigua
| `--o Holopea Hall, 1847
| |-- H. insignis
| |-- H. pyrene
| `--+-- H. rotunda
| `-- H. symmetrica
`--+--o Sinutropis Perner, 1903
| |-- Pachystrophia spiralis
| |-- Sinutropis ?esthetica
| `-- Euomphalus tubus
`--o Pachystrophia Perner, 1903
|-- P. gotlandica
`--+-- Lytospira triquestra
`-- Lytospira subuloides
Reference(s):
Wagner, P. J., 2002 [1995]: Phylogenetic relationships of the earliest anisostrophically coiled gastropods.
--Smithsonian Contributions to Paleobiology: Vol. 88, pp. vi-152
[Author notes that scientific material of this article is based on his 1995 Ph.D.
dissertation, which is partly outdated]
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checked ATW090318
cladogram and footnote © Mikko Haraamo
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