Molluscomorpha - Proto-molluscs
Status
The molluscomorpha are the (largely theoretical) line
between Lophotrochozoa and Mollusca. Nearly everyone's best guess at
present is that the Halkeriida are very close to that line. At the
moment, we cover nothing which actually falls between Halkeriida and
Mollusca, so this entry is essentially a placeholder, eventually to be
filled by Kimberella, Hyolitha, and probably sipunculid
worms. A cladogram -- no more speculative than most -- is included
below.
Our working cladogram looks like this:
Lophotrochozoa
|--Halkeriida
`--+--BRACHIOPODA
`--MOLLUSCOMORPHA
|
MOLLUSCA
|
Halkieria evangelista
length about 3.7 cm
stratigraphic range: Atdabanian Age (other Halkeriid species known from the early Tommotian)
horizon: Sirius Passet formation, Greenland
illustration from Jerzy Dzik, "Early Metazoan Evolution and the Meaning of It's Fossil Record", fig. 11 p.367
Tentative Molluscomorphan Cladogram
This cladogram combines Conway Morris 1993,
Conway Morris 1998, and Runnegar 2002. Both the Order Halkeriida and the Class Coeloscleritophora are considered
paraphyletic grades.
<==o LOPHOTROCHOZOA
`--o COELOSCLERITOPHORA (paraphyletic)
|--+-- †Chancelloriida
| `-- †Sachitida
`--+-- †Siphogonuchitida
`--o †Halkieriida (paraphyletic)
|-- †Halkeria evangelista
|--o "MOLLUSCOMORPHA"
| |?- †Kimberella
| `--+?- †Tullimonsterida
| |--o Mollusca
| | `--+-- SOLENOGASTRES
| | `--+-- †Acaenoplax
| | |-- CAUDOFOVEATA
| | `--+-- TESTARIA
| | `--+-- POLYPLACOPHORA
| | `-- CONCHIFERA
| `--+-- †Hyolitha
| `-- Sipuncula
|--o "ANNELIDAMORPHA"
| |?- †Wiwaxia corrugata
| `--+?- †Canadia spinosa
| `-- Annelida
`--o LOPHOPHORATA

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page uploaded 30 November 2002
checked ATW031215
page by
M. Alan Kazlev
unless otherwise indicated by copyright notice, this material may be freely used for non-commercial purposes
