Palæos:  Palaeos Home Page Procoelomates
INVERTEBRATES Molluscomorpha

Page Back Unit Back Up Unit Home
Page Next Unit Next Down

Molluscomorpha - Proto-molluscs

Status

The molluscomorpha are the (largely theoretical) line between Lophotrochozoa and Mollusca.  Nearly everyone's best guess at present is that the Halkeriida are very close to that line.  At the moment, we cover nothing which actually falls between Halkeriida and Mollusca, so this entry is essentially a placeholder, eventually to be filled by Kimberella, Hyolitha, and probably sipunculid worms.  A cladogram -- no more speculative than most -- is included below.

Our working cladogram looks like this:

Lophotrochozoa
|--Halkeriida
`--+--BRACHIOPODA
   `--MOLLUSCOMORPHA
      |
      MOLLUSCA
 

click for larger image
Halkieria evangelista
length about 3.7 cm


stratigraphic range: Atdabanian Age (other Halkeriid species known from the early Tommotian)
horizon: Sirius Passet formation, Greenland

illustration from Jerzy Dzik, "Early Metazoan Evolution and the Meaning of It's Fossil Record", fig. 11 p.367

Tentative Molluscomorphan Cladogram

This cladogram combines Conway Morris 1993, Conway Morris 1998, and Runnegar 2002. Both the Order Halkeriida and the Class Coeloscleritophora are considered paraphyletic grades.

<==o LOPHOTROCHOZOA
   `--o COELOSCLERITOPHORA (paraphyletic)
      |--+-- †Chancelloriida
      |  `-- †Sachitida
      `--+-- †Siphogonuchitida
         `--o †Halkieriida (paraphyletic)
            |-- †Halkeria evangelista
            |--o "MOLLUSCOMORPHA"
            |  |?- †Kimberella
            |  `--+?- †Tullimonsterida
            |     |--o Mollusca
            |     |  `--+-- SOLENOGASTRES
            |     |     `--+-- †Acaenoplax
            |     |        |-- CAUDOFOVEATA
            |     |        `--+-- TESTARIA
            |     |           `--+-- POLYPLACOPHORA 
            |     |              `-- CONCHIFERA
            |     `--+-- †Hyolitha
            |        `-- Sipuncula
            |--o "ANNELIDAMORPHA"
            |  |?- †Wiwaxia corrugata
            |  `--+?- †Canadia spinosa
            |           `-- Annelida 
            `--o LOPHOPHORATA

Page Back Unit Home Page Up Page Top Page Next



images not loading? | error messages? | broken links? | suggestions? | criticism?
contact webmaster

page uploaded 30 November 2002
checked ATW031215
page by M. Alan Kazlev
unless otherwise indicated by copyright notice, this material may be freely used for non-commercial purposes