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Glossary R-S


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Rhizoid: hair-like filamentous projection for anchorage or absorption.  

Rhizome: a (usually) underground stem that is horizontally oriented; rhizomes may appear like roots, but have a definite node and internode architecture. Image from the site: Flora of Roosevelt Monmouth County, New Jersey by Ross Tulloss & Mike Hamilton.

Root: a plant organ that functions in anchorage and absorption; in seed plants derived from a bipolar embryo

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Saccate pollen: pollen with a saccus or sacci; characteristic of many conifers.  See image at bisaccate

Saccus: ( pl. sacci) a wing-like or bladder-like extension on a pollen grain.  See image at bisaccate.

Sclerenchyma: tissue composed of cells with walls thickened with lignin; sclerenchyma tissue functions primarily in strengthening and support 

Rhynchosperma seed Dunn et al. (202)Sclerotesta: the middle, fibrous layer of the integument in some seeds.  Image from Dunn et al. (2002) showing cross sections of a Rhynchosperma quinii seed from the late Mississippian (Serpukhovian?) of Arkansas.

Secondary growth: growth in width initiated and maintained by the vascular cambium and cork cambium.  See cambium.  The vascular cambium surrounds the core of the stem, branch or root.  It creates xylem medially and phloem radially.  If present, the cork cambium, a second, outer layer of cambium, creates a new outer layer (typically bark) radially.  

Secondary phloem: phloem produced by the vascular cambium. See phloem, cambium, secondary growth.

Secondary xylem xylem produced by the vascular cambium. See xylem, cambium, secondary growth.   

Seed: a fertilized ovule; megasporangium that contains an embryo enclosed in an integument.  A seed has been described as "a baby sporophyte in a jacket with a lunch sack."  Gymnosperm Evolution.  The advantage of a seed is that it can combine the dispersal functions of a spore with sexual reproduction.

Neoropteris (seed fern)Seed fern: any of a number of extinct seed-bearing plants with fern-like leaves. Image of Neuropteris from the Kentucky Paleontological Society web site.

Seed plants: a monophyletic clade of plants that reproduces by seeds; megagametophyte is retained on the parent sporophyte and enclosed in an integument; microgametophyte is transferred to the megagametophyte.

Sepal: a whorl of a flower; sepals often resemble reduced leaves and function in the protection of the bud; sepals may be modified to function more like a petal

Seta:  This incredibly over-used anatomical term is simply Latin for a bristle or a stiff hair.  It is used in a variety of contexts, not only for things that look like bristles (or, to be sure, stiff hairs), but also for things like the stalk of the sporophyte in mosses, which looks nothing like a bristle (or, for that matter, a stiff hair).   

Simple tissue: tissue composed of only one cell type 

Sinus: the indentations of a lobed pinna or pinnule of a fern or of a dicot leaf.

Solute: anything dissolved in a solvent.  In salt water, the water is the solvent, and the salts are solutes.

SorusSorus: (pl sori) a cluster or group of sporangia of a fern; most frequently applied to clusters of fern sporangia attached to the bottom of fern leaves. Image from the General Biology Lab site by Diane Jokinen and Patrick Duffie of Loyola University Chicago.

Sporangiophore: modified branch on which the sporangium is attached. 

Sporangium: (pl. sporangia) hollow, unicellular or pluricellular structure in which spores are produced. 

Spore: a haploid (1N) reproductive cell capable of developing directly into a gametophyte without uniting with another cell. 

Sporophyll: a spore-bearing leaf; a modified leaf bearing sporangia

Sporophyte: the spore producing, diploid (2N) phase of the life cycle; compare to gametophyte

SSU rDNA: DNA coding for RNA associated with the small ribosomal subunit, i.e. for "18S" RNA.  

Stamen: the "male" structures of a flower, consisting of a supporting filament and the anther, a sac which produces and holds the microgametophytes (pollen).  

Stele: vascular tissue (the xylem and phloem together); the central vascular cylinder in stems and roots where the vascular tissue is located. 

Stem: same as axis; a plant axis with leaves or enations

Stigma: (pl. stigmata) the distal part of the pistil, the organ that receives the pollen.  Stigmata may be quite varied in shape and structure.  They are often button-shaped and studded with hair-like papillae ("dry" stigmata) or coated with a sticky film ("wet" stigmata) to capture pollen.  

StomaStoma: (pl. stomata) tiny pores or opening in the epidermis of leaves; stomata are flanked by two guard cells that regulate opening and closing of the pore and thus regulate gas exchange (respiration) and transpiration.  Image from the Ecotree site by Martin Cocks of the University of the Western Cape (South Africa).

Strobilus: (pl. strobili) a cone a cone-shaped cluster of spore-bearing leaves (e.g., a pine cone), an aggregation of sporophylls on a common axis

Suberin: waxy, waterproof chemical in some plant cells, notably cork (in stems) and endodermis cells (in roots).  Suberin is an extremely complex and irregular material, like lignin -- with which it shares some similarities.  Suberin is composed of two physically separated domains: the aliphatic and phenolic.  The phenolic domain is rather lignin-like, but with even greater variability, and built on the same basic unit of a di- or tri-hydroxyphenyl group attached to a three-carbon chain, variously oxidized and integrated with the carbohydrates of the cell walls.  Perhaps the most common building block is ferulic acid: formally, 3-(2'-methoxy-3'-hydroxyphenyl)-propenoic acid.  Distally, the phenolic domain is attached at points by ester Suberin Bernards (2002) linkages to glycerol.  The remaining hydroxyls of the glycerol molecule are ester-linked to some strange-looking C-18 to C-30 lipids.  These lipids are substituted at C9-10 with one or two hydroxyls, or even with an epoxide link between the two carbons.  Finally the ω- position may be oxidized to a carboxylate (alone or esterized to glycerol) or hydroxyl (alone or esterised to ferulic acid).  Variations allow for cross-linkage to other suberin molecules via the 9-10 or ω positions. Image adapted from Bernards (2002).  

Style: a tubular column in the pistil of a flower, through which the pollen is transported from the stigmata to the ovary.

Succulent: a plant that is able to store water in its tissues and then withdraw it during times of drought. e.g. a cactus.  The water storage tissue may be found in the stem, leaves, or roots depending on the species.  Stem succulents, leaf succulents, and root succulents are types of growth forms. 

Synangium: (pl. synangia) a reproductive unit composed of fused sporangia.

Synapomorphy: a unique character which is shared by all basal members of a clade and is derived from their common ancestor, but not shared with ancestral groups. A synapomorphy may be secondarily lost in later descendants. Only a synapomorphy may be used to infer phylogeny.



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