Palaeos Plants: Lycopods: Lycopod Classification and Phylogeny

    Class Lycopsida ("club-mosses')
        Order Drepanophycales
            Family Drepanophycaceae
        Order Protolepidodendrales
            Family Protolepidodendraceae
            Family Eleutherophyllaceae
        Order Lycopodiales
            Family Lycopodiaceae
        Order Selaginellales
            Family Selaginellaceae
        Order Lepidocarpales
            Family Cyclostigmaceae
            Family Flemingitaceae
            Family Sigillariostrobaceae
            Family Lepidocarpaceae
            Family Spenceritaceae    
            Family Caudatocarpaceae
            Family Pinakodendraceae
            Family Sporangiostrobaceae
            Family Pleuromeiaceae        
        Order Miadesmiales 
            Family Miadesmiaceae
        Order Isoetales ("quillworts')
            Family Isoetaceae
            Family Chaloneriaceae

Palaeozoic Palaeobotany of Great Britain as well as listing British sites and the history of discovery of plant fossils in Britian, this is an excellent intro to Palaeobotany in general.

<==o LYCOPODIOPHYTA 
   |--+--o †ZOSTEROPHYLLOPSIDA
   |  |  |-- †Zosterophyllum
   |  |  |-- †Gosslingia
   |  |  |-- †Rebuchia
   |  |  |-- †Bathurstia
   |  |  |-- †Crenaticaulis
   |  |  |-- †Serrulacaulis
   |  |  |--o †Sawdonia
   |  |  |  |-- †S. ornata
   |  |  |  `-- †S. acanthotheca
   |  |  |-- †Discalis longistipa
   |  |  |-- †Drepanophycus spinaeformis
   |  |  |-- †Kaulangiophyton akantha
   |  |  `-- †Asteroxylon mackiei
   |  `--+--o †ASTEROXYLALES
   |     `--+--o †DREPANOPHYCALES
   |        `--+?- †Baragwanathia longifolia
   |           |?- †Archaeosigillaria
   |           `--o LYCOPODIOPSIDA)
   |              `--o LYCOPODIALES
   |                 `--o Lycopodiaceae 
   |                    |--o Huperzia 
   |                    |  `-- H. selago
   |                    |--o Lycopodiella
   |                    |  `-- L. inundata 
   |                    |--o Lycopodium 
   |                    |  |-- L. annotinum
   |                    |  `-- L. clavatum 
   |                    `--o Diphasiastrum
   |                       |-- D. alpinum 
   |                       `-- D. complanatum
   `--+--o †PROTOLEPIDODENDRALES
      |  |--o †Prolepidodendron
      |  |  `-- †P. scharyanum
      |  |--o †Estinnophyton 
      |  |  |-- †E. wahnbachense
      |  |  `-- †E. gracilis
      |  |-- †Sugambrophyton
      |  |-- †Colpodexylon
      |  |-- †Leclercqia complexa
      |  `--o †Spencerites 
      |     |-- †S. moorei
      |     `-- †S. insignis
      `--+?-o SELAGINELLALES 
         |  `--o Selaginellaceae 
         |     |-- †Selagibellites hallei
         |     |-- †Miadesmia
         |     `--o Selaginella 
         |        |-- †S. harrisiana
         |        `-- S. selaginoides 
         `--o ISOETOPSIDA 
            |--o LEPIDODENDRALES 
            |  |-- †Valmeyerodendron triangularifolium
            |  |-- †Cyclostigma
            |  |-- †Bisporangiostrobus
            |  |-- †Lepidosigillaria
            |  |--o †Sigillariaceae
            |  |  `--o †Sigillaria
            |  |     |-- †(Eusigillaria)
            |  |     |-- †(Subsigillaria)
            |  |     `-- †(Sigillaria)
            |  `--o †Lepidodendraceae
            |     |--o †Lepidodendron
            |     |  |-- †L. hickii
            |     |  `-- †L. aculaetum
            |     |-- †Paralycopodites brevifolius
            |     |--o †Diaphorodendron
            |     |  |-- †D. vaskulare
            |     |  |-- †D. phillipsii
            |     |  |-- †D. scleroticum
            |     |  `-- †D. dicentricum
            |     `--o †Lepidophloios
            |        `-- †L. halli
            `--o ISOETALES 
               |--o Isoetaceae [Isoëtaceae] 
               |  `--o Isoetes [Isoëtes] 
               |     |-- I. lakustris
               |     `-- I. echinospora
               |--o Pleuromeiales
               |  |-- †Pleuromeia sternbergi
               |  |-- †Nathorstiana arborea
               |  `-- †Chaloneria
               |-- †Paurodentron fraipontii
               |-- †Oxroadia
               `-- †Prostigmaria eggertiana

Hämet-Ahti, L., Suominen, J., Ulvinen, T., Uotila, P. & Vuokko, S., 1984: Retkeilykasvio. --Suomen Luonnonsuojelun Tuki Oy & Forssan Kirjapaino Oy, Forssa, 1984, 1-544

Links

Systematics of the Lycophyta, at the UCMP site has a simplified and incomplete lycopsid phylogeny -- but useful for getting oriented to the main groups. Another fine introduction, which will probably not be on the web very long, can be found at An Overview of Green Plant Phylogeny. A nice student paper with an interesting phylogeny: Brautigam, S, C Li, A Mudry, G Miller & E Prebil (2001), An analysis of the evolution of the vascular cryptogams, Lycopodophyta, Sphenophyta, and the Pteridophyta, through examination of extinct and extant genera. J. Sys. Biol. Susq. Univ. 8(4). See Brautigam et al. (2001). The ultimate collection of lycopsid links in general is, as usual, on the Links for Paleobotanists site. See Lycophyta, Introductions to both Fossil and Recent Plant Taxa. Having little to do with lycopsid phylogeny per se, but well worth reading: Bateman, RM, PR Crane, WA DiMichele, PR Kenrick, NP Rowe, T Speck & WE Stein (1998), Early evolution of land plants: Phylogeny, physiology, and ecology of the primary terrestrial radiation. Ann. Rev. Ecol. Syst. 29: 263–92 See Bateman et al. (1998). ATW050803.

Palæos:		Lycopsids
PLANT EVOLUTION		Classification & Cladogram

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Lycophyta after Cleal

Lycophyta cladogram (various sources)

Reference(s):

Links

Lycopsida Home	Drepanophycales	Protolepidodendrales	Lycopodiales	Selaginellales
Lepidodendrales	Miadesmiales	Pleuromeiales	Isoetales	Cladogram