Palæos:		Unit 340:Theropoda
The Vertebrates		Cladogram

Theropod Cladogram

DINSAUROMORPHA
|--ORNITHISCHIA
`--+--SAUROPODOMORPHA
   |
   Theropoda
   |--Herrerasauridae
   `--+--Ceratosauria
      `--Tetanurae
         |--Torvosauroidea
         `--Avetheropoda
            |--Carnosauria
            `--Coelurosauria
               |--Tyrannosauroidea
               `--Maniraptora
                  |--Dromaeosauridae
                  |
                  `--AVES            

DINOSAUROMORPHA
|
`--Theropoda ToL, MH
   |--Herrerasauridae •X 
   `--+--Ceratosauria •X Th, MH
      `--Tetanurae Th, MH
         |--Torvosauroidea •X
         `--Avetheropoda
            |--Carnosauria X Th, MH
            |  |--Cryolophosaurus •X
            |  `--+--Monolophosaurus •X
            |     `--Allosauroidea X Th
            |        |--Sinraptoridae •X
            |        `--Allosauridae X  
            |           |--Acrocanthosaurus •X
            |           `--Carcharodontosauridae •X
            `--Coelurosauria MH
               |--Compsognathidae •X
               `--+--Ornitholestes •X
                  `--+--Coelurus •X 
                     `--Maniraptoriformes
                         |--+--Ornithomimosauria •X MH
                         |  `--Tyrannosauroidea X ToL, MH, Th  
                         |     |--Eotyrannus •X  
                         |     `--Tyrannosauridae X  
                         |        |--Alectrosaurus •X  
                         |        `--Tyrannosaurinae X
                         |           |--+--Albertosaurus •X  
                         |           |  `--Gorgosaurus •X
                         |           `--Tyrannosaurini •X
                         `--Maniraptora Th
                            |--+--Oviraptorosauria MH
                            |  `--Therizinosauroidea X DC, Th
                            |     |--Beipiaosaurus •X
                            |     `--+--Alxasaurus •X
                            |        `--Therizinosauridae •X
                            `--+--+--Troodontidae •X DC, Th
                               |  `--Dromaeosauridae •X Th
                               |
                               `--AVES

Sources: The arrangement of genera within Carnosauria is unclear.  Our arrangement is consistent with Currie & Carpenter (2000).  Pisani et al. (2001) place Acrodontosaurus closer to Carcharodontosaurus than to Allosaurus.  We tend to agree, but deem Currie & Carpenter's direct study of the carnosaurs as better authority than the supertree of Pisani et al.  Fortunately, our cladogram lacks sufficient resolution for this to make much difference.  ATW060203.

checked ATW040810